2023

Shihao Dong , Gaoying Gu, Tao Lin , Ziqi Wang , Jianjun Li, Ken Tan*, James C. Nieh* (2023) An inhibitory signal associated with danger reduces honeybee dopamine levels. Current Biology, https://doi.org/10.1016/j.cub.2023.03.072

Zhengwei Wang, Xiuxian Chen, Frank Becker, Uwe Greggers, Stefan Walter, Marleen Werner, Charles R. Gallistel*, and Randolf Menzel* (2023) Honey bees infer source location from the dances of returning foragers. PNAS, 120(12), e2213068120. https://doi.org/10.1073/pnas.2213068120

Shihao Dong, Tao Lin, James C. Nieh*, Ken Tan*(2023)Social signal learning of the waggle dance in honey bees. Science, 379(6636): 1015-1018.  https://doi.org/10.1126/science.ade1702

2022

Yuren Li, Yufeng Qu, Huanli Xu* and Zhenwei Wang* (2022) A new record of sweat-sucking stingless bee, Lisotrigona carpenteri Engel 2000, from a natural savanna in southwest China. Journal of Apicultural Research, 61(5):594-597. https://doi.org/10.1080/00218839.2021.1987741

Yufeng Qu, Shijie Wang, Kai Wang and Zhenwei Wang* (2022) The newly rising meliponiculture and research on stingless bees in China-a mini review. Journal of Apicultural Research, 61(5): 730-737. https://doi.org/10.1080/00218839.2022.2104568 

Zhixiang Dong, Qihe Tang, Wanli. Li, Zhengwei Wang, Xijie Li, Chaomin Fu, Dan Li, Kai Qian, Wenli Tian, Jun Guo* (2022) Honeybee (Apis mellifera) resistance to deltamethrin exposure by modulating the gut microbiota and improving immunity. Environmental Pollution, 314, 120340. https://doi.org/10.1016/j.envpol.2022.120340 

Yufeng Qu, Xinxin Wen, Yuxin Wei and Zhenwei Wang* (2022) Biology of bamboo nesting stingless bees of southwest China. Apidologie, 53, 53. https://doi.org/10.1007/s13592-022-00962-1  

Junjun Zhang, Zhengwei Wang, Katrina Klett, Yufeng Qu, Ken Tan (2022) Higher toxin tolerance to triptolide, a terpenoid foraged by a sympatric honeybee. Journal of Insect Physiology, 137, 104358. https://doi.org/10.1016/j.jinsphys.2022.104358 

Shihao Dong, Aili Sun, Ken Tan*, James C. Nieh* (2022) Identification of giant hornet Vespa mandarinia queen sex pheromone components. Current Biology, 32(5):R211-R212. https://doi.org/10.1016/j.cub.2022.01.065 

Yanan Cheng, Ping Wen; Ken Tan*, Darrouzet Eric* (2022) Designing a sex pheromone blend for attracting the yellow-legged hornet (Vespa velutina), a pest in its native and invasive ranges worldwide. Entomologia Generalis,42(4):523-530. DOI: 10.1127/entomologia/2022/1395 

2021

Yichuuan Meng, Xianggui Mo, Tiantian He, Xinxin Wen, James C. Nieh* ,  Xinwang Yang and Ken Tan* (2021)New bioactive peptids from the venom gland of a social honet Vespa velutina. Toxicon,199:94-100.

Katrina Klett, Junjun Zhang, Yingying Zhang, Zhengwei Wang, Shihao Dong*, KenTan* (2021) The Nasonov gland pheromone as a potential source of death cue in Apis cerana. Journal of Insect Physiology, https://doi.org/10.1016/j.jinsphys.2021.104238.

Gaoying Gu, Yichuan Meng, Ken Tan, Shihao Dong* and James C. Nieh*(2021) Lethality of Honey Bee Stings to Heavy Armored Hornets. Biology, 2021, 10, 484. https://doi.org/10.3390/biology10060484. 

2020

Zhiwen Gong, Gaoying Gu, Yuan Wang, Shihao Dong, Ken Tan^*, James C. Nieh^* (2020) Floral tea polyphenols can improve honey bee memory retention and olfactory sensitivity, Journal of insect physiology, https://doi.org/10.1016/j.jinsphys.2020.104177  

Shihao Dong, Ken Tan*and James C. Nieh*（2020）Visual contagion in prey defense signals can enhance honest defense. Journal of Animal Ecology, 90:594-601.https://doi.org/10.1111/1365-2656.13390

2019

Zhengwei Wang*, Ken Tan* (2019) Honey Bee Alarm Pheromone Mediates Communication in Plant–Pollinator–Predator Interactions. Insects, 10, 366. https://doi.org/10.3390/insects10100366

Zhiwen Gong, Ken Tan,* and James C. Nieh,* (2019) Hornets possess long-lasting olfactory memories. Journal of Experimental Biology , 222, jeb200881. doi:10.1242/jeb.200881

Yuan Li, Xingchuan Jiang, Zhengwei Wang*, Junjun Zhang, Katrina Klett, Shahid Mehmood, Yufeng Qu, and Ken Tan,* (2019) Losing the Arms Race: Greater Wax Moths Sense but Ignore Bee Alarm Pheromones. Insects, 10(3), 81.  https://doi.org/10.3390/insects10030081

Shihao Dong, KenTan*, Qi Zhang, James C.Nieh*(2019) Playbacks of Asian honey bee stop signals demonstrate referential inhibitory communication. Animal Behaviour, 148:29-37. https://doi.org/10.1016/j.anbehav.2018.12.003

Shihao Dong, Ping Wen, Qi Zhang, Yuan Wang, Yanan Cheng, Ken Tan, *James C. Nieh* （2018）Olfactory eavesdropping of predator alarm pheromone by sympatric but not allopatric prey. Animal Behaviour, 141:115-125. doi.org/10.1016/j.anbehav.2018.05.013

Zhiwen Gong, Ken Tan* and James C. Nieh* (2018) First demonstration of olfactory learning and long term memory in honey bee queens. Journal of Experimental Biology, 221, jeb 177303.

Gao Chen*, Zhengwei Wang , Ping Wen, Wei Wei, Ya Chen, Hui Ai, Weibang Sun (2018) Hydrocarbons mediate seed dispersal: a new mechanism of vespicochory. The New Phytologist, doi: 10.1111/nph.15166

Junjun Zhang, Zhengwei Wang*, Ping Wen, Yufeng Qu, Ken Tan* and James C. Nieh (2018) The reluctant visitor: an alkaloid in toxic nectar can reduce olfactory learning and memory in Asian honey bees. Journal of Experimental Biology, 221, jeb168344. doi:10.1242/jeb.168344

2017

Ken Tan*, Cao Wang, Shihao Dong, Xinyu Li & James C. Nieh* (2017) The pesticide flupyradifurone impairs olfactory learning in Asian honey bees (Apis cerana) exposed as larvae or as adults. Scientific Reports 7, 17772

Gao Chen*, ZhengWei Wang, Yan Qin, WeiBang Sun (2017) Seed dispersal by hornets: An unusual insect-plant mutualism. Journal of Integrative Plant Biology, (on line)

Ping Wen*, Yanan Cheng, Shihao Dong, ZhengWei Wang, Ken Tan, James Nieh* (2017) The sex pheromone of a globally invasive honey bee predator, the Asian eusocial hornet, Vespa velutina. Scientific Reports, 12956

Ping Wen*, Yanan Cheng, Yufeng Qu, Hongxia Zhang, Jianjun Li, Heather Bell, Ken Tan, James Nieh (2017) Foragers of sympatric Asian honey bee species intercept competitor signals by avoiding benzyl acetate from Apis cerana alarm pheromone. Scientific Reports, 7(1), 6721

Xiao-Lan Wen, Ping Wen*, Cecilia A. L. Dahlsj？, David Sillam-Dussès, Jan ？obotník (2017) Breaking the cipher: ant eavesdropping on the variational trail pheromone of its termite prey, Proceedings of the Royal Society B: Biological Sciences, 284:1853

Isobel Ronai, Michael H. Allsopp, Ken Tan, Shihao Dong, Xiwen Liu, Vanina Vergoz, Benjamin P. Oldroyd (2017) The dynamic association between ovariole loss and sterility in adult honeybee workers. Proc. R. Soc. B 284: 20162693. http://dx.doi.org/10.1098/rspb.2016.2693

Zhiwen Gong, Chao Wang, Shihao Dong, Xuewen Zhang, Yanhui Wang, Zongwen Hu, Ken Tan*(2017) High Concentrations of the Alarm Pheromone Component, Isopentyl Acetate, Reduces Foraging and Dancing in Apis mellifera Ligustica and Apis cerana Cerana. Journal of Insect Behavior, 30(2):188-189

R. Gloag*, Ken Tan, Yuchong Wang, Wenfei Song, Weitin Luo, G. Buchman，Madeleine Beekman，Benjamin P. Oldroyd（2017）No evidence of queen thelytoky following interspecific crosses of the honey bees Apis ceranaand Apis mellifera. Insectes Sociaux, (on line), doi:10.1007/s00040-016-0538-3

Shihao Dong, Ping Wen, Qi Zhang, Xinyu Li, Ken Tan* & James Nieh (2017) Resisting majesty: Apis cerana, has lower antennal sensitivity and decreased attraction to queen mandibular pheromone than Apis mellifera. Scientific Reports 7: 44640 ,doi:10.1038/srep44640

Yanan Cheng, Ping Wen*, Shihao Dong, Ken Tan*, James C. Nieh (2017) Poison and alarm: the Asian hornet Vespa velutina uses sting venom volatiles as an alarm pheromone. Journal of Experimental Biology, 220(4): 645-651. doi: 10.1242/jeb.14878

2016

Zhiwen Gong, Chao Wang, James C. Nieh, Ken Tan* (2016) Inhibiting DNA methylation alters olfactory extinction but not acquisition learning in Apis cerana and Apis mellifera. Journal of Insect Physiology, 90:43-48. doi:10.1016/j.jinsphys.2016.05.007

Jonathan L. Teichroew , Jianchu Xu, Antje Ahrends, Zachary Y. Huang, Ken Tan, Zhenghua Xie(2016) Is China’s unparalleled and understudied bee diversity at risk? Biological Conservation (online). doi:10.1016/j.biocon.2016.05.023

Zhengwei Wang,  Ping Wen, Yufeng Qu, Shihao Dong,  Jianjun Li, Ken Tan* & James C. Nieh*（2016）Bees eavesdrop upon informative and persistent signal compounds in alarm pheromones. Scientific Reports 6：25693. doi:10.1038/srep25693

Ken Tan*, Shihao Dong, Li X, Xiwen Liu, Chao Wang, Jianjun Li & James C. Nieh* (2016) Honey Bee Inhibitory Signaling Is Tuned to Threat Severity and Can Act as a Colony Alarm Signal. PLoS Biology, 14(3): e1002423. doi:10.1371/journal.pbio.1002423

Zhengwei Wang, Yufeng Qu, Shihao Dong, Ping Wen, Jianjun Li, Ken Tan*, Randolf Menzel (2016) Honey Bees Modulate Their Olfactory Learning in the Presence of Hornet Predators and Alarm Component. PLoS ONE,11(2): e0150399. doi:10.1371/journal.pone.0150399

2015

Ken Tan, Yufeng Qu , Zhengwei Wang, Zhiwei Liu, Michael S. Engel (2015) Haplotype diversity and genetic similarity among populations of the Eastern honey bee from Himalaya-Southwest China and Nepal (Hymenoptera: Apidae). Apidologie, 47(2):197-205.

Ping Wen, Jianchu Mo, Chunwen Lu, Ken Tan, Jan ？obotník, David Sillam-Dussès (2015) Sex-pairing pheromone of Ancistrotermes dimorphus(Isoptera: Macrotermitinae). Journal of Insect Physiology, 83(1):8-14. doi:10.1016/j.jinsphys.2015.11.006

Ken Tan, Tanya Latty, Shihao Dong, Xiwen Liu, Chao Wang & Benjamin P. Oldroyd (2015) Individual honey bee (Apis cerana) foragers adjust their fuel load to match variability in forage reward. Scientific Reports, 5:16418. doi:10.1038/srep16418

Ken Tan, Yuchong Wang, Shihai Dong, Xiwen Liu, Di Zhuang, Weiwen Chen, Benjamin P. Oldroyd (2015) Associations between reproduction and work in workers of the Asian hive bee Apis cerana. Journal of Insect Physiology, 82(1):33-37. doi:10.1016/j.jinsphys.2015.08.003.

Ken Tan，Weiwen Chen, Shihao Dong, Xiwen Liu, Yuchong Wang & James C. Nieh （2015） A neonicotinoid impairs olfactory learning in Asian honey bees (Apis cerana) exposed as larvae or as adults. Scientific Reports, 5:10989 . DOI: 10.1038/srep10989

Ken Tan, Xiwen Liu, Sihao Dong, Chao Wang, Benjamin P. Oldroyd (2015) Pheromones affecting ovary activation and ovariole loss in the Asian honey bee Apis cerana. Journal of Insect Physiology, 74(1):25-29.

Michael J. Holmes, Ken Tan, Zhengwei Wang, Benjamin P. Oldroyd, Madeleine Beekman (2015) Genetic reincarnation of workers as queens in the Eastern honeybee Apis cerana. Heredity ,114 (1):65-68.  doi:10.1038/hdy.2014.70.

Ken Tan, Shihao Dong, Xiwen Liu,Weiweng Chen, Yuchong Wang,Benjamin P. Oldroyd , Tanya Latty (2015) Phantom alternatives influence food preferences in the eastern honeybee Apis cerana. Journal of Animal Ecology, 84(2):509-517. doi: 10.1111/1365-2656.12288.

2014

Jianjun Li, Zhengwei Wang, Ken Tan*, Yufeng Qu, James C. Nieh (2014) Effects of natural and synthetic alarm pheromone and individual pheromone components on foraging behavior of the giant Asian honey bee, Apis dorsata. The Journal of Experimental Biology, 217(19):3512-3518. doi: 10.1242/jeb.110171.

Jianjun Li, Zhengwei Wang, Ken Tan*, Yufeng Qu, James C. Nieh (2014) Giant Asian honeybees use olfactory eavesdropping to detect and avoid ant predators. Animal Behaviour, 97:69-76. doi: 10.1016/j.anbehav.2014.08.015.

Michael J. Holmes, Ken Tan, Zhengwei Wang, Benjamin P. Oldroyd, Madeleine Beekman (2014)Effect of queen excluders on ovary activation in workers of the Eastern honeybee Apis cerana. Insectes Sociaux, 61(2):191-196. doi:10.1007/s00040-014-0344-8.

Ken Tan*, Weiwen Chen, Shihao Dong, Xiwen Liu, Yuchong Wang, James C. Nieh (2014) Imidacloprid Alters Foraging and Decreases Bee Avoidance of Predators. PLoS ONE,9(7): e102725. doi: 10.1371/journal.pone.0102725

Michael J. Holmes, Ken Tan, Zhengwei Wang, Benjamin P. Oldroyd, Madeleine Beekman (2014) Why acquiesce? Worker reproductive parasitism in the Eastern honeybee (Apis cerana). Journal of Evolutionary Biology, 27(5):939-949. doi: 10.1111/jeb.12366.

Emily J. Remnant, Anna Koetz, Ken Tan, Eloise Hinson, Madeleine Beekman and Benjamin P. Oldroyd (2014) Reproductive interference between honeybee species in artificial sympatry. Molecular Ecology, 23(5): 1096–1107. doi: 10.1111/mec.12669.

Ken Tan, Tanya Latty, Zongwen Hu, Zhengwei Wang, Shuang Yang, Weiweng Chen, Benjamin P. Oldroyd (2014) Preferences and tradeoffs in nectar temperature and nectar concentration in the Asian hive bee Apis cerana. Behavioral Ecology and Sociobiology, 68(1):13-20. doi: 10.1007/s00265-013-1617-3.

Zhengwei Wang, Gao Chen, Ken Tan* (2014) Both olfactory and visual cues promote the hornet Vespa velutina to locate its honeybee prey Apis cerana. Insectes Sociaux, 61(1):67-70. doi:10.1007/s00040-013-0326-2.

Zhengwei Wang, Ken Tan* (2014) Comparative Analysis Of Olfactory Learning Of Apis Cerana And Apis Mellifera. Apidologie,45(1):45-52. doi:10.1007/s13592-013-0228-3.

2013

Ken Tan, Zongwen Hu, Weiwen Chen, Zhengwei Wang, Yuchong Wang, James C. Nieh (2013) Fearful Foragers: Honey Bees Tune Colony and Individual Foraging to Multi-Predator Presence and Food Quality. PLoS ONE, 8(9): e75841. doi:10.1371/journal.pone.0075841.

Ken Tan, Shuang Yang, Zhengwei Wang, Menzel R (2013) Effect of Flumethrin on Survival and Olfactory Learning in Honeybees. PLoS ONE 8(6): e66295. doi:10.1371/journal.pone.0066295.

Michael J. Holmes, Ken Tan, Zhengwei Wang, Benjamin P. Oldroyd, Madeleine Beekman (2013) Honeybee (Apis cerana) guards do not discriminate between robbers and reproductive parasites. Insectes Sociaux, 60(2):265-271. doi:10.1007/s00040-013-0292-8.

Ken Tan, Zhengwei Wang, Weiwen Chen, Zongwen Hu, Benjamin P. Oldroyd (2013) The ‘I see you’ prey–predator signal of Apis cerana is innate. Naturwissenschaften, 100(3):245-248.

2012-2010

Ken Tan, Zhengwei Wang, Mingxian Yang, Stefan Fuchs, Linjuan Luo, Zuyun Zhang, Hua Li, Di Zhuang, Shuang Yang, Juergen Tautz, Madeleine Beekman, Benjamin P. Oldroyd (2012) Asian hive bees, Apis cerana, modulate dance communication in response to nectar toxicity and demand. Animal Behaviour, 84(6):1589-1594.

Ken Tan, Shuang Yang, Zhengwei Wang, Sarah E. Radloff, Benjamin P. Oldroyd (2012) Differences in foraging and broodnest temperature in the honey bees Apis cerana and A. mellifera. Apidologie, 43(6): 618-623.

Ken Tan, Zhenwei Wang, Hua Li, Shuang Yang, Zongwen Hu, Gerald Kastberger, Benjamin P. Oldroyd (2012) An ‘I see you’ prey–predator signal between the Asian honeybee, Apis cerana, and the hornet, Vespa velutina. Animal Behaviour, 83(4):879-882.

Ken Tan, Mingxian Yang, Zhengwei Wang, Sarah E. Radloff, Christian W. W. Pirk (2012) The pheromones of laying workers in two honeybee sister species: Apis cerana and Apis mellifera. Journal of Comparative Physiology A, 198(4):319-323.

Ken Tan, Mingxian Yang, Zhengwei Wang, Hua Li, Zuyun Zhang, Sarah E. Radloff, H. Randall Hepburn(2012) Cooperative wasp-killing by mixed-species colonies of honeybees, Apis cerana and Apis mellifera. Apidologie, 43(2):195-200.

Mingxian Yang,  Sarah E. Radlof, Ken Tan, H. Randall Hepburn(2010) Anti-predator Fan-blowing in Guard Bees, Apis mellifera capensis Esch. Journal of Insect Behaviour, 23(1):12–18.

Mingxian Yang, Zhengwei Wang, Hua Li, Zuyun Zhang, Ken Tan, Sarah E. Radloff, H. Randall Hepburn (2010) Thermoregulation in mixed-species colonies of honeybees (Apis cerana and Apis mellifera). Journal of Insect Physiology, 56(7): 706-709.

Mingxian Yang, Ken Tan, Sarah E. Radlof , Mananya Phiancharoen, H. Randall Hepburn (2010) Comb construction in mixed-species colonies of honeybees, Apis cerana and Apis mellifera. The Journal of Experimental of Biology, 213, 1659-1664.

Ken Tan, Hua Li, Mingxian Yang, H. Randall Hepburn, Sarah E. Radloff (2010) Wasp hawking induces endothermic heat production in guard bees. Journal of Insect Science, 10:142.

Ken Tan, Zhengwei Wang, Mingxian Yang, H. Randall Hepburn, Sarah E. Radloff (2010) Nestmate Recognition Differences between Honeybee Colonies of Apis cerana and Apis mellifera. Journal of insect behavior, 23(5):381-388.

Ken Tan, Zhengwei Wang, Hua Li, Mingxian Yang, Christian W. W. Pirk, H. Randall Hepburn, Sarah E. Radloff (2010) Responses of Queenright and Queenless Workers of Apis cerana to 9-keto-2(E)-decenoic Acid, a Pheromonal Constituent of the Mandibular Gland. Journal of Chemical Ecology, 36(9):966-968.

Sarah E. Radloff, Colleen Hepburn, H. Randall Hepburn, Stefan Fuchs, Soesilawati Hadisoesilo, Ken Tan, Michael S. Engel and Viktor Kuznetsov (2010) Population structure and classification of Apis cerana. Apidologie, 41(6): 589–601.

Mingxian Yang, Ken Tan, Sarah E. Radloff, Christian W.W. Pirk, H. Randall Hepburn (2010) Hetero-specific queen retinue behavior of worker bees in mixed-species colonies of Apis cerana and Apis mellifera. Apidologie, 41(1):54-61.

B. Dainat, Ken Tan, H. Berthoud, P. Neumann (2009) The ectoparasitic mite Tropilaelaps mercedesae (Acari, Laelapidae) as a vector of honeybee viruses. Insect Sociaux, 56 (1):40 – 43.

Ken Tan, Mingxian Yang, Sarah E. Radlof, Christian W. W. Pirk, Robin M. Crewe, Mananya Phiancharoen, Randall Hepburn, Benjamin P. Oldroyd (2009) Worker reproduction in mixed-species colonies of honey bees. Behavioural Ecology, 20(5):1106-1110.

Ken Tan, Mingxian Yang, Sarah E. Radloff, Yushen Yu, Christian W. W. Pirk, H. Randall Hepburn ( 2009) Intra- and interspecific brood recognition in pure and mixed-species honeybee colonies, Apis cerana and A. mellifera. Apidologie, 40(2):184-191.

Jianjun Li, Ken Tan, Shaoyu He (2008) Hornet predation risk: apperception and response by honey bees. Journal of apicultural research, 47(1): 174–175.

Ken Tan, Stefan Fuchs, Michael S. Engel (2008) An adventitious distal abscissa in the forewing of honey bees (Hymenoptera: Apidae: Apis). Apidologie. 39(6):674-682.

Ken Tan, Mingxian Yang,  Sarah E. Radlof, H. Randall Hepburn, Zhuyun Zhang, Linjuan Luo, Hua Li (2008) Dancing to different tunes: heterospecific deciphering of the honeybee waggle dance. Naturwissenschaften. 95(12):1165-1168.

Ken Tan, Yanhong Guo, S. W. Nicolson, Sarah E. Radloff, Qishi Song, H. Randall Hepburn (2007) Honeybee (Apis cerana) Foraging Responses to the Toxic Honey of Tripterygium hypoglaucum (Celastraceae):Changing Threshold of Nectar Acceptability, Journal of Chemical Ecology. 33(12):2209-2217.

Ken Tan, Sarah E. Radloff, Jianjun Li, H. Randall Hepburn Mingxian Yang, Lianjiang Zhang , P. Neumann (2007) Bee-hawking by the wasp, Vespa velutina, on the honeybees Apis cerana and A. mellifera. Naturwissenschaften. 94(6):469-472.

Ken Tan, Sarah E. Radloff, H. Randall Hepburn, Mingxian Yang, Lianjiang Zhang and Xian Fan (2007) Environmentally-induced developmental effects on morphometric characters of workers in Apis cerana colonies. Apidologie, 38(3):289-295.

Ken Tan, Natapot Warritand, Deborah Roan Smith (2007) Mitochondrial DNA diversity of Chinese Apis cerana. Apidologie, 38(3):238-246.

Ken Tan, Marina D. Meixner, Stefan Fuchs, Xuan Zhang, Shaoyu He, Irfan Kandemir, Walter S. Sheppard，Nikolaus Koeniger (2006) Geographic distribution of the eastern honeybee, Apis cerana (Hymenoptera: Apidae), across ecological zones in China: Morphological and molecular analyses. Systematics and Biodiversity, 4(4): 473-482.

Ken Tan , H. Randall Hepburn, Shaoyu He, Sarah E. Radloff, P. Neumann, Xiang Fang. (2006) Gigantism in honeybees: Apis cerana queens reared in mixed-species colonies. Naturwissenschaften, 93(7):315-320.

Ken Tan, H. Randall Hepburn, Sarah E. Radloff, Yusheng Yu, Yiqiu Liu, Danyin Zhou, P. Neumann (2005) Heat-balling wasps by honeybees. Naturwissenschaften, 92(10):492-495.

Sarah E. Radloff, H. Randall Hepburn, Colleen Hepburn, Stefan Fuchs, Gard W. Otis, M.M. Sein, H.L. Aung, H.T. Pham, D.Q. Tam, A.M. Nuru, Ken Tan (2004)  Multivariate morphometric analysis of of southern mainland Asia. Apidologie, 36(1):127-139.

Ting Zhou, Denis L. Anderson, Zachary Y. Huang, Shuangxiu Huang, Jun Yao, Ken Tan, Qingwen Zhang (2004) Identification of Varroa mites infesting Apis cerana and Apis mellifera in China. Apidologie, 34(6):545-554.

Ken Tan, Stefan Fuchs, Nikolaus Koeniger, Zan Ruiguang (2003) Morphological characterization of Apis cerana in the Yunnan Province of China. Apidologie, 34(6):553-562.